Gregor Mendel formulated the Law of Independent Assortment based on his classic hybridization experiments with pea plants. However, modern geneticists know that certain physical chromosome configurations violate this law.

Based on the genetic configurations shown above, which phenomenon represents a physical exception to Mendel's Law of Independent Assortment?
Codominance of alleles at a single gene locus
Autosomal linkage of genes on the same chromosome
Multiple alleles regulating a single phenotypic trait
Epistatic interaction between different gene loci