An ecologist studying a Scottish Highland pine forest models the organization of the ecosystem using the nested framework shown in the diagram below:

Let the levels be defined as follows:
Which of the following represents the correct sequence of these ecological levels, ordered from the least inclusive (smallest) to the most inclusive (largest) level?
L4→L3→L2→L1L_4 \rightarrow L_3 \rightarrow L_2 \rightarrow L_1L4→L3→L2→L1
L4→L2→L3→L1L_4 \rightarrow L_2 \rightarrow L_3 \rightarrow L_1L4→L2→L3→L1
L1→L2→L3→L4L_1 \rightarrow L_2 \rightarrow L_3 \rightarrow L_4L1→L2→L3→L4
L4→L3→L1→L2L_4 \rightarrow L_3 \rightarrow L_1 \rightarrow L_2L4→L3→L1→L2