Leaf structure: maximising photosynthesis
Photosynthesis
The process in which plants and algae use light energy, absorbed by chlorophyll in the chloroplasts, to make glucose from carbon dioxide and water, releasing oxygen as a by-product.
- Photosynthesis is the process in which light energy absorbed by chlorophyll is used to make glucose from carbon dioxide and water, releasing oxygen.
- The word equation is carbon dioxide+water→glucose+oxygen\text{carbon dioxide}+\text{water}\rightarrow\text{glucose}+\text{oxygen}carbon dioxide+water→glucose+oxygen.
- The balanced symbol equation is 6CO2+6H2O→C6H12O6+6O26\mathrm{CO_2}+6\mathrm{H_2O}\rightarrow\mathrm{C_6H_{12}O_6}+6\mathrm{O_2}6CO2+6H2O→C6H12O6+6O2.
- A broad leaf provides a large surface area for light absorption, while a thin leaf gives carbon dioxide a short diffusion distance.
Upper layers let light reach chloroplasts
Palisade mesophyll
The layer of tall cells near the top of a leaf, packed with chloroplasts to absorb the most light for photosynthesis.
- The waxy cuticle is transparent so light passes through, and waterproof so it reduces evaporation from the surface.
- The upper epidermis is thin and transparent, allowing light to reach the palisade mesophyll.
- Palisade mesophyll is a layer of tall, closely packed cells near the upper leaf surface containing many chloroplasts.
- Its position and chloroplast density allow it to absorb a high proportion of the incoming light.
Do not state that epidermal cells contain many chloroplasts; guard cells are the main epidermal exception.
Spongy mesophyll speeds gas exchange
Spongy mesophyll
A layer of loosely packed leaf cells with large air spaces that give a big surface area for water to evaporate and gases to move.
- Spongy mesophyll is a loosely packed layer with many interconnecting air spaces and moist cell surfaces.
- The air spaces provide a large internal surface area and let carbon dioxide diffuse rapidly from stomata to photosynthesising cells.
- Oxygen made in photosynthesis diffuses through the same spaces and out of the leaf.
- The lower epidermis contains many stomata, which allow gas exchange while reducing exposure to direct sunlight and air movement.

Veins supply and remove materials
- Xylem in each vascular bundle supplies water and mineral ions to leaf cells.
- Phloem carries sucrose and amino acids away from photosynthesising cells to sources and sinks elsewhere in the plant.
- A network of veins keeps cells close to transport tissue and also helps support the leaf blade.
Example: palisade cells contain many chloroplasts, so they absorb more light and can photosynthesise at a higher rate.
Explaining adaptations in exams
- Required link: state the feature and then explain how it increases light absorption, gas diffusion, water supply or sugar removal.
- Diagram labels: use waxy cuticle, upper epidermis, palisade mesophyll, spongy mesophyll, air space, lower epidermis, guard cells, stoma, xylem and phloem.
- Why is a leaf broad and thin?
- How is palisade mesophyll adapted for photosynthesis?
- How do spongy mesophyll air spaces aid gas exchange?
- What do xylem and phloem do in a leaf vein?
- Why is the upper epidermis transparent?