Which two of the following statements regarding neural, hormonal and genetic factors in aggression are TRUE? Select the two correct letters.

Option A: According to the dual-hormone hypothesis, high levels of testosterone only promote aggressive behavior when cortisol levels are also high.

Option B: The amygdala is a subcortical structure in the limbic system that evaluates the emotional importance of sensory stimuli and triggers aggressive responses.

Option C: The MAOA gene is located on the Y chromosome, which explains why aggressive behavior is more prevalent in males.

Option D: Low levels of serotonin disrupt the inhibitory regulation of the prefrontal cortex over the amygdala, increasing the likelihood of impulsive aggression.

Option E: Twin studies show that the heritability of aggressive behavior is 100%, indicating that genetic factors are solely responsible.