During an ecological survey of a saltmarsh estuary, a researcher wishes to estimate the population sizes of two species: the European glasswort (Salicornia europaea\textit{Salicornia europaea}Salicornia europaea), a sessile halophyte, and the common shore crab (Carcinus maenas\textit{Carcinus maenas}Carcinus maenas), a highly motile decapod.
Which of the following describes appropriate protocols for estimating the population sizes of these species?
- For S. europaea\textit{S. europaea}S. europaea, lay out a grid over the saltmarsh, generate random coordinates, place frame quadrats at these coordinates, and count the individuals in each to calculate a mean density.
- For C. maenas\textit{C. maenas}C. maenas, lay out a belt transect perpendicular to the shoreline, place quadrats at 5 m intervals, and count the crabs within each quadrat.
- For C. maenas\textit{C. maenas}C. maenas, capture individuals using baited traps, mark each crab with a small dab of non-toxic, water-resistant paint on its carapace, release them, and recapture a second sample after 24 hours24\text{ hours}24 hours to calculate the Lincoln Index.