A long-term ecological monitoring programme of breeding pairs of the Northern Spotted Owl (Strix occidentalis caurina\textit{Strix occidentalis caurina}Strix occidentalis caurina) in a coniferous forest reserve was carried out over several decades. In 1995, the number of breeding pairs recorded was 5,850. In 2022, the number of breeding pairs recorded was 1,480.
Calculate the mean annual decrease in the population of individual spotted owls in this reserve between 1995 and 2022. Assume each breeding pair consists of two individual birds. Show your working.
Give your answer to three significant figures and include appropriate units.
